Highlighted: where the king and rook start and land.
Weirdest castling #11
SP-14 · qnnrkbbr
O-O · K e1→g1 · R h1→f1
54 of the 960 starts castle exactly this way.
Destinations are fixed in Chess960 (kingside K→g1 R→f1, queenside K→c1 R→d1); only the starting squares differ.